2-Down’s boss

This crossword clue is for the definition: 2-Down’s boss.
it’s A 13 letters crossword puzzle definition.
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Possible Answers: MEL.

Last seen on: LA Times Crossword 5 Nov 2017, Sunday

Random information on the term “MEL”:

The mouse erythroleukemia cell line F4N(MEL) is a mouse dervied aneuploid cell line. It was established in 1977. Eight percent of the total number of cells in culture undergo spontaneous apoptosis.

MEL on Wikipedia