Ruler in ‘War and Peace’

This crossword clue is for the definition: Ruler in ‘War and Peace’.
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Possible Answers: TSAR.

Last seen on: USA Today Crossword – Jan 10 2021

Random information on the term “TSAR”:

Stramenopile is a taxonomic concept that identifies a clade of organisms. A clade is conceived as any group of all of the descendants of a common ancestor (i.e. it is both holophyletic and monophyletic). A clade can be defined by reference to an evolutionarily innovative feature that the ancestor and its descendents share acknowledging that the character may be secondarily lost. The stramenopile clade is distinguished by the presence of stiff tripartite external hairs. In most species, the ahirs are attached to flagella, in some they to other parts of the surface of cells, and in some they have been secondarily lost (in which relatedness to stramenopile ancestors is evident from other shared cytological features or from similarity in the makeup of genes).

Members of the clade are referred to as ‘stramenopiles’. Most stramenopiles are single celled eukaryotes (protists), but stramenopiles include some multicellular algae and fungi. The group includes a variety of algal protists, heterotrophic flagellates, opalines and closely related proteromonad flagellates (all endobionts in other organisms); the actinophryid heliozoa, and oomycetes. The tripartite hairs have been lost in some stramenopiles – for example in most diatoms (although they still make the proteins – see below).

TSAR on Wikipedia